The answer: http://antcat.org/documents/6914/Scha%C ... graphy.pdf
Schär, S.; Talavera, G.; Espadaler, X.; Rana, J. D.; Andersen, A. A.; Cover, S. P.; Vila, R. 2018. Do Holarctic ant species exist? Trans-Beringian dispersal and homoplasy in the Formicidae. Journal of Biogeography 00:1-12. [2018-06-28]