The loss of flight in ant workers enabled an evolutionary redesign of the thorax for ground labour.
Christian Peeters, Roberto A. Keller, Adam Khalife, Georg Fischer, Julian Katzke, Alexander Blanke, Evan P. Economo.
https://link.springer.com/article/10.1186/s12983-020-00375-9
or
https://link.springer.com/cont…86/s12983-020-00375-9.pdf with 9 supplemental files.